Juan Armando Sánchez, Ph.D.





Sánchez J.A., Aguilar C, Dorado D, Manrique N. 2007. Phenotypic plasticity and morphological integration in a marine modular invertebrate. BMC Evolutionary Biology 7(1): 12.


Sánchez J.A.  2007. A new genus of Atlantic octocorals (Octocorallia: Gorgoniidae): systematics of gorgoniids with asymmetric sclerites. Journal of Natural History 41(9-12): 493-509.


Grajales A, Aguilar C, Sanchez JA. 2007 Phylogenetic reconstruction using secondary structures of Internal Transcribed Spacer 2 (ITS2, rDNA): finding the molecular and morphological gap in Caribbean gorgonian corals. BMC Evolutionary Biology 7: 90.


Aguilar C, & Sánchez JA. 2007. Phylogenetic hypotheses of gorgoniid octocorals according to ITS2 and their predicted RNA secondary structures. Molecular Phylogenetics & Evolution 43(3): 774-86. doi:10.1016/j.ympev.2006.11.005


Tracey, M D., Neil, H., Marriott, P., Andrews, H A., Cailliet, M G., Sanchez, J.A. 2007. Age and growth of two genera of deep-sea bamboo corals (family isididae) in New Zealand waters. Bulletin of Marine Science 81(3): 393-408.


Aguilar C, & Sánchez JA. 2007. Molecular morphometrics: contribution of ITS2 sequences and predicted RNA secondary structures to octocoral systematics. Bulletin of Marine Science




Mcfadden C. S., S. C. France, J.A. Sánchez & P. Alderslade. 2006. A molecular phylogenetic analysis of the Octocorallia (Cnidaria: Anthozoa) based on mitochondrial protein-coding sequences. Molecular Phylogenetics and Evolution, 41(3), 2006, 513-527


Grajales A. & Sánchez JA. 2006. Discharged nematocysts of Millepora alcicornis. Coral Reefs. 25: 671.



Ward, J.R., Rypien, K., Bruno, J., Harvell, C.D., Jordan, E., Mullen, K., Rodriguez-Martinez, R., Sanchez, J.A & Smith, G. Coral Diversity and Disease in Mexico. Diseases of Aquatic Organisms 69: 23-31


Sánchez J.A.  & A.A. Rowden.  2006 Octocoral Diversity on New Zealand Seamounts. Proceedings of the 10th International Coral Reef Symposium Japan 5:




Sánchez J.A. 2005. Systematics of the bubblegum corals (Paragorgiidae: Octocorallia: Cnidaria) with description of new species from New Zealand and the Eastern Pacific. Zootaxa 1014: 1-72.


Opresko D.M. & J.A. Sánchez. 2005. Caribbean Shallow-water Black Corals (Cnidaria: Anthozoa: Antipatharia). Caribbean Journal of Science 41(3): 492 – 507.


Sánchez J. A. &H. Wirshing. 2005. A field key to the identification of zooxanthellate octocorals from the Caribbean and Western Atlantic. Caribbean Journal of Science 41(3): 508 – 522.


Collin R., M. C. Díaz, J.Norenburg, R. M. Rocha, J. A. Sánchez, A. Schulze, M. Schwartz, & A. Valdés. 2005. Photographic Identification Guide to Some Common Marine Invertebrates of Bocas Del Toro, Panama. Caribbean Journal of Science 41(3): 638-707.


Sánchez J.A.  , V. Pizarro, A. R. Acosta, P. A. Castillo, P. Herron, J. C. Martínez, P. Montoya & C. Orozco. 2005 Evaluating Coral Reef benthic Communities from remote Caribbean Atolls (Quitasueño, Serrana, and Roncador Banks, Colombia): multivariate approaches to recommend Marine Protected Areas for the SeaFlower Biosphere Reserve (Archipelago of San Andres and Providencia). Atoll Research Bulletin 531: 1-65.


Garzon S. P., A. D. Rodriguez, J. A. Sanchez & Eduardo Ortega-Barria, 2005. Sesquiterpenoid metabolites with antiplasmodial activity from a Caribbean gorgonian coral, Eunicea sp. Journal of Natural Products, 68, 1354-1359.


Ospina, C.A., Rodriguez, A.D., Sanchez, J.A., Ortega-Barria, E., Capson, T.L., And Mayer, A.M.S. 2005 Caucanolides a-f, unusual antiplasmodial constituents from a Colombian collection of the gorgonian coral Pseudopterogorgia bipinnata. Journal of Natural Products 68,(10): 1519 – 1526.




Sánchez J.A., H.R.Lasker, E. G. Nepomuceno, J. D. Sánchez & M. J. Woldenberg. 2004. Branching and Self-Organization in Marine Modular Colonial Organisms: a Model. The American Naturalist 163 (3) (0003-0147/2004/16303-20458).


Sánchez J.A.  2004. Evolution and dynamics of branching colonial form in marine modular Cnidarians: Gorgonian Octocorals. Hydrobiologia. 530 (1): 283-290.

 (uncorected proof)


Sánchez J.A., M.F. Gil, L.H. Chasqui & E.M. Alvarado. 2004. Grazing dynamics on a Caribbean reef-building coral. Coral Reefs 23(4): 578-583. (DOI: 10.1007/s00338-004-0419-5)


Sánchez J.A., & S.D. CAIRNS. 2004. An unusual new gorgonian coral from the Aleutian Islands, North Pacific, Alaska. Zoologishe Mededelingen- Leiden 78: 265-274.

uncorrected proof


Charry, H. E.M. Alvarado & Sánchez J.A. 2004. Annual skeletal extension of two reef-building corals from the Colombian Caribbean Sea. Boletín de Investigaciones Marinas y Costeras 33: 209-222.


Wei, X., Rodriguez A.D., Bran P., Raptis R.G., Sánchez, J.A., E. Ortega-Barria & J. Gonzales. 2004. Antiplasmoidal cembradiene diterpenoids from a Southwestern Caribbean gorgonian octocoral of the genus Eunicea. Tetrahedron 60: 11813-11819.


Rodriguez, I., Shi Y-P., Garcia O.J., Rodriguez A.D., Mayer A.M.S., Sánchez, J.A., E. Ortega-Barria & J. Gonzales. 2004. New Pseudopterosin and seco-Pseudopterosin diterpene glycosides from two Colombian isolates of Pseudopterogorgia elisabethae and their diverse biological activities. Journal of Natural Products 67(10): 1672-1680.


Marrero J, Rodriguez Ad, Baran P, Raptis Rg, Sanchez J.A, Ortega-Barria E, Capson Tl. 2004. Bielschowskysin, a gorgonian-derived biologically active diterpene with an unprecedented carbon skeleton. Organic Letters 6(10):1661-4.





Sánchez J. A., C. S. Mcfadden, S. C. France, & H. R. Lasker. 2003. Molecular Phylogenetic analyses of shallow-water Caribbean octocorals. Marine Biology 142: 975-987. (DOI 10.1007/s00227-003-1018-7)


Sánchez J.A., H.R.Lasker & D.J. Taylor. 2003. Phylogenetic analyses among octocorals (Cnidaria) according to mitochondrial and nuclear DNA sequences (lsu-rRNA 16S, and ssu-rRNA 18S) support two convergent clades of branching gorgonians. Molecular Phylogenetics & Evolution. 29(1): 31-42. (DOI:10.1016/S1055-7903(03)00090-3)


Sánchez J.A., & H.R.Lasker. 2003. Patterns of Morphologic Integration in Branching Colonies of Marine Modular Organisms: supra-module organization in Gorgonian Corals. Proceedings of the Royal Society of London B. 270: 2039-2044. (DOI: 10.1098/rspb.2003.2471)


Sánchez J.A., & H.R.Lasker. 2003. Do multi-branched colonial organisms exceed normal growth after partial mortality? Proceedings of the Royal Society of London B (suppl.). (DOI 10.1098/rsbl.2003.0708)


Sánchez J.A., W. Zeng, V. R. Coluci, C. Simpson & H. R. Lasker. 2003. How similar are branching networks in nature? A view from the ocean: Caribbean gorgonian corals. Journal of Theoretical Biology. 222: 135-138. (DOI:10.1016/S0022-5193(03)00017-1)


Lasker H.R., M. L. Boller, J. Castanaro & Sánchez J.A. 2003. Modularity and determinate growth in a gorgonian coral. The Biological Bulletin 205: 319-330.


Friedlander A., Sladek-Nowlis J., Sanchez J. A., Appeldoorn R., Usseglio P., Mccormick C., Bejarano S., And Mitchell-Chui A. 2003. Designing Effective Marine Protected Areas in Old Providence and Santa Catalina Islands, San Andrés Archipelago, Colombia, Using Biological and Sociological Information. Conservation Biology 17(6): 1769-1784.




Sánchez J.A. 2002. Branching and evolution of modular colonial organisms (Ph.D. thesis). State University of New York at Buffalo.


Lasker H.R. & Sánchez J.A. 2002. Allometry and Astogeny of modular organisms. pp. 207-253. In: R.N. Hughes (ed.), Reproductive Biology of Invertebrates. Vol. XI. Progress in Asexual Reproduction. John Wiley, New York.


Shi, Y.-P.; Rodríguez, A.D; Barnes, C.L.; Sánchez, J.A.; Raptis, R.G.; Baran, P. 2002. New Terpenoid Constituents from Eunicea pinta. Journal of Natural Products 65 (9):1232-1241. (DOI: 10.1021/np0200820




Lasker H.R. & Sánchez J.A. 2001. Octocorals: growth in two and three dimensions.  In: J. Kaandorp & J. Klueber (eds.) The Algorithmic Beauty of Seaweeds, Sponges and Corals. Springer-Verlag, Amsterdam. cover


Sánchez J.A. 2001. Systematics of the Southwestern Caribbean Muriceopsis Aurivillius (Cnidaria: Octocorallia: Gorgonacea) with description of a new species. Bulletin of the Biological Society of Washington. 10: 160-180.




Diaz Jm., Diaz-Pulido G. & Sánchez J.A.  2000. Distribution and structure of the southernmost Caribbean coral reefs: Golfo de Urabá, Colombia. Scientia Marina. 64(3): 327-336.




Sánchez J.A. 1999. Black coral-octocoral distribution patterns on a deep-water reef, Imelda bank, Caribbean sea, Colombia. Bulletin of Marine Science. 65(1): 215-225.


Sánchez J.A., E.M. Alvarado, M.F. Gil, O. Arenas, H. Charry, L.H. Chasqui & R.P. Garcia. 1999. Synchronous mass spawning of Montastraea annularis (Ellis & Solander) and M. faveolata (Ellis & Solander) (Faviidae: Scleractinia) at Rosario islands, Caribbean coast of Colombia. Bulletin of Marine Science. 65(3): 873-879.




Sánchez J.A., J.M. Diaz &. S. Zea. 1998. Octocoral and black coral distribution patterns on the barrier reef-complex of Providencia island, Southwestern Caribbean. Caribbean Journal of Science. 34 (3-4): 250-264.




Sánchez J.A., S. Zea &. J.M. Diaz. 1997. Gorgonian communities of two contrasting environments from oceanic Caribbean atolls. Bulletin of Marine Science. 61(2): 61-72.


Diaz, J.M., Sánchez J.A. & J. Geister. 1997. Development of Lagoonal reefs in oceanic complexes of the Southwestern Caribbean: geomorphology, structure and distribution. Proceedings of the VIII International Coral Reef Symposium, Panama 1: 779-784.


Cobar, O., Rodríguez A.D., Padilla O.L. & J.A. Sánchez 1997. The Calyculaglycosides: Dilophol-type diterpene glycosides exhibiting antiinflammatory activity from the Caribbean gorgonian Eunicea sp. The Journal of Organic Chemistry 62 (21): 7183-7188.



Diaz, J.M., Sánchez J.A., J. Garzon-Ferreira & S. Zea. 1996. Morphology and marine habitats from two southwestern Caribbean atolls: Albuquerque and Courtown. Atoll Research Bulletin 435: 1-33.


Diaz, J.M., Sánchez J.A.& G. Diaz. 1996. Morfología y formaciones coralinas recientes de Isla Fuerte y Bajo Bushnell, plataforma continental del Caribe colombiano. Boletín de Investigaciones Marinas y Costeras. 25: 87-105.


Diaz, J.M., G. Diaz, J. Garzon-Ferreira, J. Geister, Sánchez J.A. & S. Zea. 1996. Atlas de los arrecifes coralinos del Caribe colombiano. I. Archipiélago de San Andrés y Providencia. Publicaciones Especiales del INVEMAR 75p.




Sánchez J.A. 1995. Benthic communities and geomorphology of the Tesoro Island reefs, Colombian Caribbean. Anales del Instituto de Investigaciones Marinas de Punta Betín.24: 55-77.



Sánchez J.A. & G.R. Navas. 1994. Notes on the distribution, habitat and morphology of Stylaster roseus (Stylasterina: Hidrozoa) at the Colombian Caribbean. Anales del Instituto de Investigaciones Marinas de Punta Betín 23: 193-197.


Sánchez J.A. 1994. Presencia de Stylatula diadema Bayer (Pennatulacea) y Carijoa riisei (Duchassaing & Michelotti) (Telestacea) en el Caribe suroccidental, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betín 23: 137-147.


For recent (2008-2009) publications please visit my lab page at: